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The plot is a continuation of the Resistance: Fall of Man storyline. The game starts with a Black Ops aircraft taking Sgt. Nathan Hale to the base of the Special Projects Research Administration (SPRA) in Iceland. Hale arrives at the base and discovers that there are others like him that are resistant to the Chimera virus who have been collected together from around the world by the SPRA. The result is a task force called 'The Sentinels'. The base then comes under attack by Chimera forces and 'The Sentinels' manage to escape to the United States. Two years after the Chimeran defeat in Great Britain, the Chimera launch a full-scale aerial invasion on both the east and west coasts of the United States